The genus Hadrolaelia re-groups, as we have already seen, the species before placed in the genus Laelia, sections Hadrolaelia Schlecther and Crispae Pftizer, as well some species of Sophronitis. It is sure that this genus is not, and the results obtained by van den Berg e all (op. cit.) confirmed this, perfectly homogenous, on the contrary, it is structured in distinct sub-groups: the figure 2, which reproduces part of the figure 6 (page109) by the authors, specifies this structure.
If the shared characters of those species, and mentioned above, seem to us to be enough to re-group in only one genus, we also believe that it is preferable to distinguish several groups. It seems evident to us to conserve the sections Hadrolaelia and Crispae, even though the exact boundary need to be better delimitated; and to create a section including the uniflora species of Sophronitis, it is useful to remember that, according to the results of van den Berg et al. those last would not form a monophyletic group.



 
Fig. 2 Cladogramme of the genus Hadrolaelia (adapted from van den Berg et al. 2.000)

  Section Crispae
  3 species, in general, placed in this section, present, however, morphological characters that bring us to separate them. Those 3 species (Hadrolaelia perrinii , Hadrolaelia virens and Hadrolaelia fidelensis) occupy particular positions, in the phylogenetic tree obtained by van den Berg et al.

 
Photo:: Guy Chiron
Hadrolaelia perrinii
Photo: Vitorino P. Castro Neto
Hadrolaelia virens
Photo: Vitorino P. Castro Neto
Hadrolaelia fidelensis

  Brieger (1981) used to separate Laelia perrinii from the other species of section Crispae, but the criterions he used in his key to determinate the sections are not very convincing. Withner (1990) keeps this section but excluding Brieger's criterions and evoking the pointed lip of this species. However, according to Pabst & Dungs (1975), the lip of the Laelia perrinii is not more pointed than the lip of Laelia crispa. The analysis of Hadrolaelia perrinii shows, in spite of all this, features that are distinctive to it: new growths protected by membranous bracts which are not tightly surrounding them but, quite the contrary, they keep away as they are inflated; the reddish reverse of the leaves ; swollen ovary, unusual column, arched, with undulated posterior face, tapering from the base towards the apex. As, to finish, this species seems, according to the results obtained by van den Berg et al. to clearly remain at a distance from the other species of the section, we propose to maintain (4) the section Perriniae by Withner.

Hadrolaelia fidelensis, though close to the other members of the section, shows remarkable differences: smooth and relatively small pseudobulbs when compared with the leaves, the very long flower stalk haste floral, small sheath, small flowers (twice smaller than the flowers of Hadrolaelia purpurata), very pointed petals, very short column. Fowlie (1977) used to say that this species "seems, regarding to the vegetative plan, a species of the section Cattleyodes, but small as a species of the section Hadrolaelia". By the time of the discovering, people put forth the idea if it was not a natural hybrid between two species of those two sections. The long flower stalk and the general feature of the flower remind, on the other hand, Laelia anceps. It seems natural to us to put it into a distinct section, intermediary between the sections Hadrolaelia and Crispae. We notice that the results obtained by van den Berg et al. seem to support this conception. Below, we propose to name this section Fidelensis.

Another species, Hadrolaelia virens, also shows peculiar features that justify a particular approach: much smaller flowers than the flowers of the other species of the section, plants coming from higher and cooler regions (1000-1200m). Those features seem to be intermediate between those from the section Crispae and those from the section Hadrolaelia. However, this species is very different from Hadrolaelia fidelensis (more elongate pseudobulbs versus rounded -peduncle shorter than the leave versus higher - little opened flower versus spread- elongate column versus short) and it does not seem suitable to us to place it inside the same section. Here, we propose to place it inside a distinct section named Virens.


(4) An alternative solution would have been to move this section to the status of sub-section inside the section Crispae.
  Section Hadrolaelia
 

Withner (1990) did not include Laelia sincorana in the conception of this section carrying on the argument about the presence of the lateral lobes in the lip of this species, which would not exist in other species of the section. Quite the contrary, our own observations, as well as Pabst & Dungst's drawings (1975), show that the big part of the species of this group has lateral lobes, more or less prominent (except for Hadrolaelia praestans, which lip is just slight widening in this level). Thus, we do not keep the section Sincoranae by Withner.

Brief, we propose the organization of the genus Hadrolaelia into six sections.

   

  (5)
  Diagnosis - Pseudobulbi cum 1-2 internodis; inflorescentia uniflora, interdumbiflora, non ortha ex spatha; pedunculus folio brevior, sine nodo; labelum longitudinaliter cristatum.

Synonymy:
Laelia Lindley section Hadrolaelia Schlecther 1917
Laelia Lindley sub-section Lobatae Cogniaux 1901. in part.


 (5) Antonyme

 
  basionym: Laelia Lindley section Crispae Pfitzer 1889, in Orchidaceae, in Engler & Prantl, Die Naturlichen Pflanzenfamilien, V.2 (6): 147.
Pfitzer defines his section in this way: 'size of the Cattleya do grupo Mossia, labelo lip slightly trilobate. Laelia crispa Lindl., Laelia purpurata Lindl., both from Brazil.
In order to define more precisely our section, we propose the follow diagnose:
 
Photo: Vitorino P. Castro Neto

Diagnosis - Pseudobuli cum multis internodis; inflorescentia multiflora, orta ex spatha ; pedunculus folio brevior, cum nodis ; labellum sine crista longitudinali, sed interdum prominente venosum.

Synonymy:
Laelia Lindley section Cattleyodes Schlecther 1917

Type species : Hadrolaelia crispa (Lindley) Chiron & V. P. Castro in Richardiana II (1): 16.2002
basionym: Cattleya crispa Lindley 1828, in Bot. Reg. 14 t. 1172

 
 

Diagnosis - Pseudobulbi cum multi internodis ; inflorescentia multiflora, orta ex spatha ; pedunculus folio brevior, cum nodis ; flores ad 5 cm diametro.

Type species: Hadrolaelia virens (Lindley) Chiron & V.P. Castro in Richardiana II (1): 17. 2002
basionym: Laelia virens Lindley 1844, in Edward's Bot. Reg. 30: misc 1.


 
 

Diagnosis - Pseudobulbi rotundari, foliis multo breviores ; rhizoma elongatum : inflorescentia foliis longior ; flores mediocres ; petala labellum que spicula ; columna brevis.

Type species : Hadrolaelia fidelensis (Pabst) Chiron & V. P. Castro in Richardiana II (1): 17. 2002
basionym: Laelia fidelensis Pabst 1967, in Orquídea 29 :11.

 
  basionym: Laelia Lindley section Perriniae Withner 1990, in The Cattleyas and their Relatives - Vol. 2 - The Laelias, p. 18.

Withner (1990) defines his section in this way: unifoliate plant with big flowers ; slightly trilobate lip; elongate sigmoid column, relatively concave in the front, tapering in the apex."

Type species : Hadrolaelia perrinii (Lindley) Chiron & V. P. Castro in Richardiana II (1): 17. 2002
basionym: Cattleya perrinii Lindley, 1838, in Edwards's Bot. 24: t.2

 
 
Photo:: Guy Chiron

Diagnosis - Pseudobulbi cum solo internodo ; inflorescentia uniflora, raro biflora, non orta & spatha ; pedunculus folio longior, sine nodo ; labellum sine crista nec venosum ; columna brevis, cum 2 grandis lateralibus alis.

Synonymy:
Eunannos Porto & Brade 1935, in Arq. Inst. Biol. Veg., Vol 2, p. 212

Type species : Hadrolaelia coccinea (Lindley) Chiron & V. P. Castro in Richardiana II (1): 17. 2002
basionym: Cattleya coccinea Lindley, 1836, in Edwards's Bot. Reg. 22: sub t. 1919.

 

Artificial key to determinate genera and sections.
1.
Plants from Central America, epiphyte, with ovoide-piriforme pseudobulbs, with inflorescence clearly higher than leaves; petals clearly larger than sepals
Laelia
  Brazilian plants ; do not present all those characters at the same time
2

2. Bifoliate rampant plants, with fusiform pseudobulbs
Microlaelia
  unifoliate plants, different feature
3

3. Rupicolous plants with claviforme spikes, inflorescence much higher or higher than leaves
Hoffmannseggella
  Epiphytic plants, inflorescence with peduncle often shorter than leaf (except for section Sophronitis)
4

4. Cylindrical, thin stems ; inflorescence without sheath, much more shorter than leaves, small flowers (<5cm), similar sepals and petals, lip re-curved towards the base and back
Dungsia
  Stems more or less thin but always more or less swollen ; big flowers in general (except for H. virens), petals larger than sepals, lip non re-curved towards the base and back
(Hadrolaelia) 5

5.		 Pseudobulb with only one internode, uniflora sometimes biflora inflorescence without a sheath
6
  Pseudobulb with many internodes, in general, multiflora inflorescence developing from a sheath
7

6. Peduncle shorter than leaf, lip with keels section Hadrolaelia
  Peduncle higher than the leaf, lip without keel section Sophronitis

7. Flowers of 5cm diameter, maximum, not widely opened section Virens
  Wide opened flowers, more than 5cm diameter 8

8. Pseudobulb much shorter than leaf section Fidelensis
  Pseudobulb and leaf with similar length 9

9. Bract of the new shoot growing clearly aside from it section Perriniae
  Bract of the new shoot growing close to it section Crispae


 
   
 

Certain species that belong to the genera above naturally hybridize between themselves as well as with the species of the genus Cattleya. A hybrid with a species of Encyclia has been also described.
We propose bellow the appropriated nothogenera.

x Hadrocattleya V. P. Castro & Chiron, in Richardiana II (1): 19. 2002
(Cattleya Lindley x Hadrolaelia (Schlechter) Chiron & V. P. Castro)

x Hadrodungsia V. P. Castro & Chiron, in Richardiana II (1): 19. 2002
(Dungsia Chiron & V.P. Castro x Hadrolaelia (Schlechter) Chiron & V. P. Castro)

x Hoffmanncattleya V. P. Castro & Chiron, in Richardiana II (1): 19. 2002
(Cattleya Lindley x Hoffmannseggella H.G. Jones)

x Hoffmanncyclia Chiron & V. P. Castro, in Richardiana II (1): 19. 2002
(Encyclia Hooker x Hoffmannseggella H.G. Jones)

x Microcattleya V. P. Castro & Chiron, in Richardiana II (1): 19. 2002
(Cattleya Lindley x Microlaelia [Schlechter] Chiron & V. P. Castro)

 
   
 

The list of the species here maintained is based mainly on Withner (1990), Miranda (1993, 1999) and Zappi (1995).The list of the species here maintained is based mainly on Withner (1990), Miranda (1993, 1999) and Zappi (1995).

Dungsia

Dungsia brevicaulis (H.G. Jones) Chiron & V. P. Castro, in Richardiana II (1): 19. 2002
basionym: Hoffmannseggella brevicaulis H. G. Jones 1972, in Rhodora 74:284

Dungsia harpophylla (Rchb.f.) Chiron & V. P. Castro, in Richardiana II (1): 19. 2002
cf. supra.
synonym: Laelia geraensis Barb. Rodr. 1877

Dungsia kautskyi (Pabst) Chiron & V. P. Castro, in Richardiana II (1): 19. 2002
basionym: Laelia kautskyi Pabst 1970, in Orch. Rev. 78:321


Hadrolaelia section Crispae

Hadrolaelia crispa (Lindley) Chiron & V. P. Castro, in Richardiana II (1): 19. 2002

Hadrolaelia grandis (Lindley & Paxton) Chiron & V. P. Castro, in Richardiana II (1): 19. 2002
basionym: Laelia grandis Lindley & Paxton 1850-1851, in Paxt. Fl. Gard. 1:60

Hadrolaelia lobata (Lindley) Chiron & V. P. Castro, in Richardiana II (1): 19. 2002
basionym: Cattleya lobata Lindley 1848, in Gard. Chron. 403
synonym: Laelia boothiana Rchb. f. 1855 - Laelia rivieri Carr. 1874

Hadrolaelia purpurata (Lindely & Paxton) Chiron & V. P. Castro, in Richardiana II (1): 19. 2002
basionym: Laelia purpurata Lindley & Paxton 1852, in Paxt. Fl. Gard. 3:111.
synonyms: Laelia casperiana Rchb. f. 1859 - Laelia russelliana hort. ex B. S. Williams 1885

Hadrolaelia tenebrosa (Rolfe) Chiron & V. P. Castro, in Richardiana II (1): 19. 2002
basionym: Laelia grandis Lindley & Paxton var. tenebrosa Rolfe 1891, in Gard Chron. ser. 3, 10:126
synonym: Laelia tenebrosa (Rolfe) Rolfe 1893.

Hadrolaelia xanthina (Lindley) Chiron & V. P. Castro, in Richardiana II (1): 19. 2002
basionym: Laelia xanthina Lindley 1859, in Bot. Mag. 85: t. 5144
synonym: Laelia wetmorei Ruschi 1970



Hadrolaelia section Fidelensis

Hadrolaelia fidelensis (Pabst) Chiron & V. P. Castro, in Richardiana II (1): 17. 2002

Hadrolaelia section Hadrolaelia

Hadrolaelia alaorii (Brieger & Bicalho) Chiron & V. P. Castro, in Richardiana II (1): 20. 2002
basionym: Laelia alaorii Briger & Bicalho 1976, in Bradea 1(17):108

Hadrolaelia dayana (Rchb. f.) Chiron & V. P. Castro, in Richardiana II (1): 20. 2002
basionym: Laelia dayana Rchb. f. 1876, in Gard. Chron. n.s. 2:772.

Hadrolaelia jongheana (Rchb. f.) Chiron & V. P. Castro, in Richardiana II (1): 20. 2002
basionym: Laelia jongheana Rchb. f. 1872, in Gard. Chron. 425.

Hadrolaelia praestans (Linden & Rchb. f.) Chiron & V. P. Castro, in Richardiana II (1): 20. 2002
basionym: Laelia praestans Linden & Rchb. f. 1857, in Allgemeine Gartenzeitung 25:336
synonym: Cattleya spectabilis Paxton 1850

Hadrolaelia pumila (Hooker) Chiron & V. P. Castro, in Richardiana II (1): 11. 2002

Hadrolaelia sincorana (Schltr.) Chiron & V. P. Castro, in Richardiana II (1): 20. 2002
basionym: Laelia sincorana Schltr.1917, in Orchis 11:72.


Hadrolaelia section Perriniae

Hadrolaelia perrinii (Lindley) Chiron & V. P. Castro, in Richardiana II (1): 17. 2002

Hadrolaelia section Sophronitis

Hadrolaelia acuensis (Fowlie) Chiron & V. P. Castro, in Richardiana II (1): 20. 2002
basionym: Sophronitis acuensis Fowlie 1975,, in Orchid Digest 39(3):150.

Hadrolaelia bicolor (Miranda) Chiron & V. P. Castro, in Richardiana II (1): 20. 2002
basionym: Sophronitis bicolor Miranda 1991, in Die Orchidee 42(5): 227

Hadrolaelia brevipedunculata (Cogn.) Chiron & V. P. Castro, in Richardiana II (1): 21. 2002
basionym: Sophronitis wittigiana Barb. Rodr. var. brevipedunculata Cogn. 1898-1902, in Martius' Flora Brasiliensis, vol 3, partie 5:318.
synonym: Sophronitis brevipedunculata (Cogn.) Fowlie 1972.

Hadrolaelia coccinea (Lindley) Chiron & V. P. Castro, in Richardiana II (1): 17. 2002
synonym: Sophronitis grandiflora Lindley 1838 - Sophronitis militaris Rchb. f. 1862
Sophronitis rossiteriana Barb. Rodr. 1877.

Hadrolaelia mantiqueirae (Fowlie) Chiron & V. P. Castro, in Richardiana II (1): 21. 2002
basionym: Sophronitis coccinea (Lindley) Rchb. f. ssp. mantiqueirae Fowlie 1968, in Orchid Digest 32(9):272-273.
synonym: Sophronitis mantiqueirae (Fowlie) Fowlie 1972.

Hadrolaelia pygmaea (Pabst) Chiron & V. P. Castro, in Richardiana II (1): 21. 2002
basionym: Sophronitis coccinea (Lindley) Rchb. f. subsp. pygmaea Pabst 1976, in Bradea 2(12)70
synonym: Sophronitis pygmaea (Pabst) Withner.


Hadrolaelia wittigiana (Barb. Rodr.) Chiron & V. P. Castro, in Richardiana II (1): 21. 2002
basionym: Sophronitis wittigiana Barb. Rodr. 1878, in Revue Horticole: 24
synonym: Sophronitis purpurea Rchb. f. 1878 - Sophronitis rosea Hort. ex Gasting 1884 -
Sophronitis violacea Lindley 1840.

Hadrolaelia section Virens

Hadrolaelia virens (Lindley) Chiron & V. P. Castro, in Richardiana II (1): 17. 2002
synonym: Laelia goebeliana Kupper & Kraenzl. 1916 - Laelia johniana Schltr. 1912

Hoffmannseggella

Hoffmannseggella alvaroana (F.E.L. Miranda) V. P. Castro & Chiron, in Richardiana II (1): 21. 2002
basionym: Laelia alvaroana F.E. L. Miranda 1999 Bradea 8(22):123.

Hoffmannseggella angereri (Pabst) V. P. Castro & Chiron, in Richardiana II (1): 21. 2002
basionym: Laelia angereri Pabst 1975, in Orch.Dig. 39:153.

Hoffmannseggella bahiensis (Schltr.) H. G. Jones 1974 in Hawai Orchid J. 3(4):16.

Hoffmannseggella blumenscheinii (Pabst) V. P. Castro & Chiron, in Richardiana II (1): 21. 2002
basionym: Laelia blumenscheinii Pabst 1975, in Bradea 1(50):487.

Hoffmannseggella bradei (Pabst) V. P. Castro & Chiron, in Richardiana II (1): 21. 2002
basionym: Laelia bradei (Pabst) 1973, in Bradea 1(31):332.

Hoffmannseggella briegeri (Blumensch. ex Pabst) V. P. Castro & Chiron, in Richardiana II (1): 22. 2002
basionym: Laelia briegeri Blumensch. ex Pabst 1973, in Bradea 1(36):367.

Hoffmannseggella cardimii (Pabst & Mello) V. P. Castro & Chiron, in Richardiana II (1): 22. 2002
basionym: Laelia cardimii Pabst & Mello 1978, in Bradea 2 (33):225-230.

Hoffmannseggella caulescens (Lindley)H.G. Jones 1970 in Caldasia 10:943.
synonym: Laelia crispilabia A. Rich ex Warner 1865 - Laelia lawranceana Warner 1865 - Laelia mantiqueirae Pabst ex Zappi 1995.

Hoffmannseggella cinnabarina (Batem. ex Lindley) H.G. Jones 1968 in Acta Bot. Acad. Sci. Hungar 14:69.

Hoffmannseggella crispata (Thunb.) H.G. Jones 1974 in Hawaii Orchid J. 3 (4):15.
synonym: Laelia cowani Rolfe 1900 - Laelia flava Lindley 1839 - Laelia fulva Lindley ex Heynh. 1841 - Laelia gardneri Pabst ex Zappi 1995.

Hoffmannseggella duveenii (Fowlie) V. P. Castro & Chiron, in Richardiana II (1): 22. 2002
basionym: Laelia duveenii Fowlie 1988, in Orch. Dig. 51:180.

Hoffmannseggella endsfeldzii (Pabst) V. P. Castro & Chiron, in Richardiana II (1): 22. 2002
basionym: Laelia endsfeldzii Pabst 1975, in Bradea 2(10):51.

Hoffmannseggella esalqueana (Blummensch. ex Pabst) V. P. Castro & Chiron, in Richardiana II (1): 22. 2002
basionym: Laelia esalqueana Blummensch. ex Pabst 1973, in Bradea 1(36):367.

Hoffmannseggella fournieri (Cogniaux) V. P. Castro & Chiron, in Richardiana II (1): 22. 2002
basionym: Laelia longipes Rchb. f. var. fournieri Cogn. 1897, in Chronique Orchidéenne 1:27.
synonym: Laelia fournieri E.F.L. Miranda 1993.

Hoffmannseggella ghillanyi (Pabst) H.G. Jones 1974 in Hawaii Orchid J. 3
(4):16

Hoffmannseggella gloedeniana (Hoehne) V. P. Castro & Chiron, in Richardiana II (1): 22. 2002
basionym: Laelia gloedeniana Hoehne 1933 (1934), in Bol. Agric. S. Paulo 34:624.

Hoffmannseggella gracilis (Pabst) V. P. Castro & Chiron, in Richardiana II (1): 22. 2002
basionym: Laelia gracilis Pabst 1979, in Bradea 2 (48):314

Hoffmannseggella hispudula (Pabst & Mello) V. P. Castro & Chiron, in Richardiana II (1): 22. 2002
basionym: Laelia hispudula Pabst & Mello 1978, in Bradea 2(33):227

Hoffmannseggella itambana (Pabst) V. P. Castro & Chiron, in Richardiana II (1): 23. 2002
basionym: Laelia itambana Pabst 1973, in Bradea 1931):333.

Hoffmannseggella kettieana (Pabst) V. P. Castro & Chiron, in Richardiana II (1): 23. 2002
basionym: Laelia kettieana Pabst 1976, in Bradea 2(22):1

Hoffmannseggella liliputana (Pabst) H. G. Jones 1974 in Hawaii Orchid J. 3(4):16.

Hoffmannseggella longipes (Rchb. f.) V. P. Castro & Chiron, in Richardiana II (1): 23. 2002
basionym: Laelia longipes Rchb. f . 1874, in Xenia Orch. 2:59.
synonym: Laelia lucasiana Rolfe 1893 - Laelia ostermayeri Hoehne ex Hoehne 1938

Hoffmannseggella macrobulbosa (Pabst) H. G. Jones 1974 in Hawaii Orchid J. 3(4):16

Hoffmannseggella milleri (Blumensch. ex Pabst) V. P. Castro & Chiron, in Richardiana II (1): 23. 2002
basionym: Laelia milleri Blumensch. ex Pabst 1973, in Bradea 1(36):367.

Hoffmannseggella mixta (Hoehne) V. P. Castro & Chiron, in Richardiana II (1): 23. 2002
basionym: Laelia mixta Hoehne 1938, in Arch. Bot. São Paulo 1:20

Hoffmannseggella munchowiana (F.E.L. Miranda) V. P. Castro & Chiron, in Richardiana II (1): 23. 2002
basionym: Laelia munchowiana F.E.L. Miranda 1999, in Bradea 8 (22):126.

Hoffmannseggella pfisteri (Pabst & Senghas) V. P. Castro & Chiron, in Richardiana II (1): 23. 2002
basionym: Laelia pfisteri Pabst & Senghas 1975, in Die Orchidee 26:254

Hoffmannseggella reginae (Pabst) V. P. Castro & Chiron, in Richardiana II (1): 23. 2002
basionym: Laelia reginae Pabst 1975, in Bradea 2:24.

Hoffmannseggella rupestris (Lindley) V. P. Castro & Chiron, in Richardiana II (1): 23. 2002
basionym: Laelia rupestris Lindley 1842, in Edwards' Bot. Reg. 28: t 62
synonym: Sophronitis mirandai van den Berg & M. W. Chase 2000

Hoffmannseggella sanguiloba (Withner) V. P. Castro & Chiron, in Richardiana II (1): 23. 2002
basionym: Laelia sanguiloba Withner 1990, in Cattleyas and their relatives, Vol. 2: The Laelias: 122

Hoffmannseggella tereticaulis (Hoehne) H.G. Jones 1974 in Florida Orchidist. 17(4):156, as"teretecaulis"

Hoffmannseggella verboonenii (F.E.L.Miranda) V. P. Castro & Chiron, in Richardiana II (1): 23. 2002
basionym: Laelia verboonenii F.E.L.Miranda 1993, in Bradea 6(18:157.


Microlaelia

Microlaelia lundii (Rchb. f.) Chiron & V.P. Castro
cf. supra
synonym: Laelia lundii Rchb. f. 1881 - Laelia reichenbachiana Wendl. & Kraenzl.1892


Natural Hybrids

Dungsia x gerhard-santosii (Pabst) Chiron & V. P. Castro, in Richardiana II (1): 24. 2002
basionym: Laelia gerhard-santosii Pabst 1974, in Bradea 1(47):472
(Dungsia harpophylla [Rchb. f.] Chiron & V. P. Castro x Dungsia kautskyi (Pabst) Chiron & V. P. Castro)

Hadrolaelia x adrienne (Rolfe) Chiron & V. P. Castro, in Richardiana II (1): 24. 2002
basionym: Laelia x adrienne Rolfe 1908, in Orchid Ver. 16:31
(Hadrolaelia jongheana [Rchb.f] Chiron & V. P. Castro x Hadrolaelia pumila [Hook.] Chiron & V. P. Castro)

Hadrolaelia x espirito-santensis (Pabst) Chiron & V. P. Castro, in Richardiana II (1): 24. 2002
basionym: Laelia x espirito-santensis Pabst 1975, in Bradea 2(4):14.
(Hadrolaelia pumila [Hook.] Chiron & V. P. Castro x Hadrolaelia xanthina [Lindley] Chiron & V. P. Castro)

Hadrolaelia x lilacina (Philbrick ex Veitch) Chiron & V. P. Castro, in Richardiana II (1): 24. 2002
basionym: Laelia x lilacina Philbrick ex Veitch 1897, in Man. Orch. Pl. 2:89.
(Hadrolaelia crispa [Lindley] Chiron & V. P. Castro x Hadrolaelia perrinii [Lindley] Chiron & V. P. Castro)

Hadrolaelia x wyattiana (Rchb.f.) Chiron & V. P. Castro, in Richardiana II (1): 24. 2002
basionym: Laelia x wyattiana Rchb.f. 1883, in Gard. Chron. II 426.
(Hadrolaelia crispa [Lindley] Chiron & V. P. Castro x Hadrolaelia lobata [Lindley] Chiron & V. P. Castro)

Hoffmannseggella x caetensis (Pabst) V. P. Castro & Chiron, in Richardiana II (1): 24. 2002
basionym: Laelia x caetensis Pabst 1975, in Bradea 2(10):50.
(Hoffmannseggella crispata [Thunb] H. G. Jones x Hoffmannseggella rupestris [Lindley] V. P. Castro & Chiron)

Hoffmannseggella x carassana (Pabst) V. P. Castro & Chiron, in Richardiana II (1): 24. 2002
basionym: Laelia x carassana Pabst 1975, in Bradea 2(6):22.
(Hoffmannseggella caulescens [Lindley] H. G. Jones x Hoffmannseggella longipes [Rchb. f.] V. P. Castro & Chiron)

Hoffmannseggella x cipoensis (Pabst) V. P. Castro & Chiron, in Richardiana II (1): 25. 2002
basionym: Laelia x cipoensis Pabst 1975, in Bradea 2(4):14.
(Hoffmannseggella ghillanyi [Pabst] H. G. Jones x Hoffmannseggella rupestris [Lindley] V. P. Castro & Chiron)

Hoffmannseggella x mucugense (F.E.L. Miranda) V. P. Castro & Chiron, in Richardiana II (1): 25. 2002
basionym: Laelia x mucugense F.E.L. Miranda 1993, in Bradea 8(18):159.
(Hoffmannseggella bahiensis [Schltr.] H. G. Jones x Hoffmannseggella pfisteri [Pabst & Senghas] V. P. Castro & Chiron)


x Hadrodungsia zaslawskii (L.C. Menezes) V. P. Castro & Chiron, in Richardiana II (1): 25. 2002
basionym: Laelia zaslawskii L.C. Menezes 1996, in Bol. Caob 24:11.
(Dungsia harpophylla [Rchb. f.] Chiron & V. P. Castro x Hadrolaelia praestans [Linden & Rchb.f.] Chiron & V. P. Castro)

x Hadrocattleya albanensis (Rolfe) V. P. Castro & Chiron, in Richardiana II (1): 25. 2002
basionym: x Laeliocattleya albanensis Rolfe 1893, in Gard. Chron. 2:584
(Cattleya warneri T. Moore x Hadrolaelia grandis [Lindley & Paxton.] Chiron & V. P. Castro)

x Hadrocattleya amanda (Rchb. f.) V. P. Castro & Chiron, in Richardiana II (1): 25. 2002
basionym: x Laelia amanda Rchb. f. 1882, in Gard. Chron. n.s., 18:776
(Cattleya intermedia Graham x Hadrolaelia lobata [Lindley] Chiron & V. P. Castro)

x Hadrocattleya amoena (Bleu) V. P. Castro & Chiron, in Richardiana II (1): 25. 2002
basionym: x Laeliocattleya amoena Bleu 1892, in Linenai 8:65, t. 367.
(Cattleya loddigesii Lindley x Hadrolaelia perrinii [Batem.] Chiron & V. P. Castro)

x Hadrocattleya binotii (Cogn.) V. P. Castro & Chiron, in Richardiana II (1): 25. 2002
basionym: x Laeliocattleya binotii Cogn. 1900, in Gard. Chron. s. 3, 28:370
(Cattleya bicolor Lindley x Hadrolaelia pumila [Hook.] Chiron & V. P. Castro)

x Hadrocattleya cypheri (Rolfe) V. P. Castro & Chiron, in Richardiana II (1): 25. 2002
basionym: x Laeliocattleya cypheri Rolfe 1897, in Orchid rev. 5:313.
(Cattleya forbesii Lindley x Hadrolaelia purpurata [Lindley & Paxton] Chiron & V. P. Castro)

x Hadrocattleya dayana (Pabst) V. P. Castro & Chiron, in Richardiana II (1): 25. 2002
basionym: x Laeliocattleya dayana Pabst 1975, in Bradea 2:68.
(Cattleya velutina Rchb. f. x Hadrolaelia crispa [Lindley] Chiron & V. P. Castro)

x Hadrocattleya delicata (Rolfe) V. P. Castro & Chiron, in Richardiana II (1): 25. 2002
basionym: x Laeliocattleya delicata Rolfe 1901, in Orchid. Rev. 9:61.
(Cattleya forbesii Lindley x Hadrolaelia crispa [Lindley] Chiron & V. P. Castro)

x Hadrocattleya elegans (Rchb. f.) V. P. Castro & Chiron, in Richardiana II (1): 26. 2002
basionym: Laelia x elegans Rchb. f., 1855, in Allg. Gartenzeitung 13;242.
(Cattleya tigrina A. Rich. ex Beer x Hadrolaelia purpurata [Lindley & Paxton] Chiron & V.P. Castro)
synonym: Laelia brysiana Lem. 1856 - Laelia devoniensis Rchb. f. - Laelia gigantea Warner 1862 - Laelia pachystele Rchb. f. - Laelia turneri Hort. ex Warner 1862

x Hadrocattleya gottoiana (Hort.) V. P. Castro & Chiron, in Richardiana II (1): 26. 2002
basionym: Laelia x gottoiana Hort. 1891, in Gard. Chron. 379
(Cattleya warneri T. Moore ex Warner x Hadrolaelia tenebrosa [Rolfe] Chiron & V. P. Castro)

x Hadrocattleya isaacii (Rolfe) V. P. Castro & Chiron, in Richardiana II (1): 26. 2002
basionym: Laeliocattleya x isaacii Rolfe 1908, in Orchid. Rev. 16:29.
(Cattleya harrisoniana Batem ex Lindley x Hadrolaelia jongheana [Rchb. f.) Chrion & V. P. Castrro ou Hadrolaelia xanthina [Lindley] Chiron & V. P . Castro)

x Hadrocattleya leeana (Rchb. f.) V. P. Castro & Chiron, in Richardiana II (1): 26. 2002
basionym: Laelia x leeana Rchb. f. 1882, in Gard. Chron. 1:492.
(Cattleya loddigesii Lindley x Hadrolaelia pumila [Hook.] Chiron & V. P . Castro)

x Hadrocattleya occhioniana (Brade) V. P. Castro & Chiron, in Richardiana II (1): 26. 2002
basionym: Laeliocattleya x occhioniana Brade 1944, in Orquidea 6:133, fig 7-12.
(Cattleya harrisoniana Batem ex Lindley x Hadrolaelia perrinii [Lindley] Chiron & V. P . Castro)

x Hadrocattleya pittiana (O'Brien) V. P. Castro & Chiron, in Richardiana II (1): 26. 2002
basionym: Laeliocattleya x pittiiana O'Brien 1894, in Gard. Chron. 1:264.
(Cattleya amethystoglossa Linden & Rchb. f. x Hadrolaelia grandis [Lindley & Paxton] Chiron & V. P . Castro)

x Hadrocattleya porphyritis (Rchb. f.) V. P. Castro & Chiron, in Richardiana II (1): 26. 2002
basionym: Laelia x porphyritis Rchb. f. 1886, in Gard. Chron. 1:72.
(Cattleya dormaniana [Rchb. f.] & Rchb. f. x Hadrolaelia pumila [Hook.] Chiron & V. P . Castro)

x Hadrocattleya reginae (Pabst.) V. P. Castro & Chiron, in Richardiana II (1): 26. 2002
basionym: Laeliocattleya x reginae Pabst 1978, in Bradea 2:280.
(Cattleya elongata Barb. Rodr. x Hadrolaelia sincorana [Schltr.] Chiron & V. P . Castro)

x Hadrocattleya rigbyana (Pabst.) V. P. Castro & Chiron, in Richardiana II (1): 27. 2002
basionym: Laeliocattleya x rigbyana Pabst 1975, in Orchid Rev. 83 (984):212
(Cattleya schilleriana Rchb. f. x Hadrolaelia pumila [Hook.] Chiron & V. P . Castro)

x Hadrocattleya schilleriana (Rchb. f.) V. P. Castro & Chiron, in Richardiana II (1): 27. 2002
basionym: Laelia x schilleriana Rchb. f. , in Otto & Dietr. Allg. Gartenz.23:323.
(Cattleya intermedia Graham x Hadrolaelia purpurata [Lindley & Paxton] Chiron & V. P . Castro)

x Hadrocattleya sgarbii (Ruschi) V. P. Castro & Chiron, in Richardiana II (1): 27. 2002
basionym: Laeliocattleya x sgarbii Ruschi 1950, in Orquidea 12:48
(Cattleya warneri T. Moore ex Warner x Hadrolaelia pumila [Hook.] Chiron & V. P . Castro)

x Hadrocattleya verelii (Rolfe) V. P. Castro & Chiron, in Richardiana II (1): 27. 2002
basionym: Laeliocattleya x verelii Rolfe 1899, in Orchid Rev. 7:340.
(Cattleya forbesii Lindley x Hadrolaelia lobata [Lindley] Chiron & V. P . Castro)


x Hoffmanncattleya lambari (Pabst) V. P. Castro & Chiron, in Richardiana II (1): 27. 2002
basionym: Laeliocattleya x lambari Pabst 1974, in Bradea l(47):472.
(Cattleya loddigesii Lindley x Hoffmmannseggella crispata [Thunb.] H. G. Jones)

x Microcattleya cattleyioides (Rich.) V. P. Castro & Chiron, in Richardiana II (1): 27. 2002
basionym: Laelia x cattleyiodes Rich 1843, in Hort. Universel 5:206.
(Cattleya loddigesii Lindley x Microlaelia lundii [Rchb. f.] Chiron & V. P . Castro)
synônimo: x Laeliocattleya fredna

x Hoffmanncyclia intermedia (F.E.L. Miranda) Chiron & V. P. Castro, in Richardiana II (1): 27. 2002
basionym: x Encylaelia intermedia F. E. L. Miranda, 1991, in Albertoa 3:94.
(Encyclia duveeni Pabst x Hoffmannseggella rupestris [Lindley.] V. P. Castro & Chiron.



Brieger, F. G.
1981
 In F. G. Brieger, R. Maatsch & K. Senghas, Schlecther´s Die Orchideen, 3a. edição, volume 1A, Verlag Paul Parey, Berlim
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A. Vogler
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 Gene trees, species trees and systematics: a cladistic perspective. Annual Review of Ecology and Systematics, 27:423-450
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1901
 Orchidaceae, in Martius' Flora Brasiliensis, p. 270-272
Fowlie, J. A.
1977
 In Brasil: part XIII - In search of the Missing Laelia fidelensis Pabst. Orch. Dig. 41(4): 147
Halbinger, F. & M. Soto
1997
 Laelias of Mexico. Orquidea (Mex.) vol. 15, número único p. 21
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 Studies in Neotropical Orchidology. Acta Botanica Academiae Scientiarum Hungaricae, T. 14 (1-2)
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 Classification phylogénétique du vivant. Belin, Paris
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 Laelia. Bot. Reg. 28: sub t.62
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Estudos em Laelia (Orchidaceae) - 1. Bradea 6 (18):155-164
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 Estudos em Laelia (Orchidaceae) - 3. Bradea 8 (22):123-127
Pabst G. F. & F. Dungs,
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Porto P. Campos & A. C. Brade
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 Arquivos do Instituto de Biologia Vegetal 2:212
Schlechter, R. von
1917
 Die Einteilung der Gattung Laelia und die geographische Verbreitung ihrer Gruppen. Orchis 11 (5):87-96
van den Berg, C. et all
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 A phylogenetic analysis of Laeliinae (Orchidaceae) based on sequence data from internal transcribed spacerts (ITS) of nuclear ribsomal DNA. Lindleyana, 15(2):96-114
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 The Cattleyas and their Relatives vol. 2: The Laelias. Timber Press, Portland, Oregon
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  Translation done by Delfina de Araujo from the French original published
in the magazine Richardiana II (1) January 2002, with authors' permission.
  Photos: Guy Chiron and Vitorino P. Castro Neto


 




(*) Guy R. Chiron
2, Rue des Pervenches,
F-38349 - VOREPPE
France		   (**) Vitorino P. Castro Neto
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